MD/PhD Graduation 2023

Alumni

Our alumni make us proud every day. Their accomplishments, achievements, and commitment to the highest quality of patient care demonstrate the not only the quality of student that this program attracts, but also the quality of physician-scientist it produces. The UK MD/PhD Program has graduated 60+ students since its inception in 1988, boasting a 98% completion rate and an over 80% long-term retention rate in academic medicine. Graduates match at top tier medical institutions in a wide range of medical specialties (see below). For many of our alumni, their education at UK made a difference in their life and is now making a difference in the lives of others. 

2021

  • Luke Heil, MD, PhD
    • Graduate Department: Microbiology, Immunology, and Molecular Genetics
    • Dissertation Title: "The Role of CD8 T Cell Immunodominance and Regulatory T Cells in Neonatal Immunity to Influenza Virus"
    • PhD Mentor: Beth A. Garvy, PhD
    • Residency: Pediatrics - University of Kentucky, Lexington, Ky.
       
  • Andrew Welleford, MD, PhD
    • Graduate Department: Anatomy and Neurobiology
    • Dissertation Title: "Autologous Peripheral Nerve Grafts to the Brain for the Treatment of Parkinson's Disease"
    • PhD Mentor: Greg Gerhardt, PhD
    • Residency: Neurology - University of Kentucky, Lexington, Ky.
       
  • Jacqueline Kulbe, MD, PhD
    • Graduate Department: Neuroscience
    • Dissertation Title: "Neuroprotective Strategies Following Experimental Traumatic Brain Injury: Lipid Peroxidation-Derived Aldehyde Scavenging and Inhibition of Mitochondrial Permeability Transition"
    • PhD Mentor: Edward D. Hall, MD
    • Residency: Psychistry/Research - UC San Diego Med Ctr, San Diego, Calif.
       
  • Scott Thalman, MD, PhD
    • Graduate Department: Biomedical Engineering
    • Dissertation Title: "Calibrated Short TR Recovery MRI for Rapid Measurement of Brain-Blood Partition Coefficient and Correction of Quantitative Cerebral Blood Flow"
    • PhD Mentor: Ai-Ling Lin, PhD
    • Residency: Med-Prelim/Radiology - U Kansas SOM, Kansas City, MO | Diagnostic Radiology - U Kansas SOM, Kansas City, Mo.
       
  • Joshua Mitchell, MD, PhD
    • Graduate Department: Molecular and Cellular Biochemistry
    • Dissertation Title: "Computational Tools for the Untargeted Assignment of FT-MS Metabolomics Datasets"
    • PhD Mentor: Hunter N. B. Moseley, PhD
    • Residency: Los Alamos National Laboratory
2021
  • Luke Heil, MD, PhD
    • Graduate Department: Microbiology, Immunology, and Molecular Genetics
    • Dissertation Title: "The Role of CD8 T Cell Immunodominance and Regulatory T Cells in Neonatal Immunity to Influenza Virus"
    • PhD Mentor: Beth A. Garvy, PhD
    • Residency: Pediatrics - University of Kentucky, Lexington, Ky.
       
  • Andrew Welleford, MD, PhD
    • Graduate Department: Anatomy and Neurobiology
    • Dissertation Title: "Autologous Peripheral Nerve Grafts to the Brain for the Treatment of Parkinson's Disease"
    • PhD Mentor: Greg Gerhardt, PhD
    • Residency: Neurology - University of Kentucky, Lexington, Ky.
       
  • Jacqueline Kulbe, MD, PhD
    • Graduate Department: Neuroscience
    • Dissertation Title: "Neuroprotective Strategies Following Experimental Traumatic Brain Injury: Lipid Peroxidation-Derived Aldehyde Scavenging and Inhibition of Mitochondrial Permeability Transition"
    • PhD Mentor: Edward D. Hall, MD
    • Residency: Psychistry/Research - UC San Diego Med Ctr, San Diego, Calif.
       
  • Scott Thalman, MD, PhD
    • Graduate Department: Biomedical Engineering
    • Dissertation Title: "Calibrated Short TR Recovery MRI for Rapid Measurement of Brain-Blood Partition Coefficient and Correction of Quantitative Cerebral Blood Flow"
    • PhD Mentor: Ai-Ling Lin, PhD
    • Residency: Med-Prelim/Radiology - U Kansas SOM, Kansas City, MO | Diagnostic Radiology - U Kansas SOM, Kansas City, Mo.
       
  • Joshua Mitchell, MD, PhD
    • Graduate Department: Molecular and Cellular Biochemistry
    • Dissertation Title: "Computational Tools for the Untargeted Assignment of FT-MS Metabolomics Datasets"
    • PhD Mentor: Hunter N. B. Moseley, PhD
    • Residency: Los Alamos National Laboratory

2020

Name Graduate Department Dissertation Title PhD Mentor Residency Evan Lynch, MD, PhD Microbiology, Immunology, and Molecular Genetics "Elucidating the Complex Signaling Events Driving Intestinal Stem Cell Plasticity Following Injury" Terrence A. Barrett, MD Plastic Surgery (Integrated) - U Kentucky Med Ctr, Lexington, KY
2020
Name Graduate Department Dissertation Title PhD Mentor Residency Evan Lynch, MD, PhD Microbiology, Immunology, and Molecular Genetics "Elucidating the Complex Signaling Events Driving Intestinal Stem Cell Plasticity Following Injury" Terrence A. Barrett, MD Plastic Surgery (Integrated) - U Kentucky Med Ctr, Lexington, KY

2019

Name Graduate Department Dissertation Title PhD Mentor Residency Kristin B. Linscott, MD, PhD Molecular and Cellular Biochemistry "Discovering a Novel Antifungal Target in Downstream Sterol Biosynthesis Using a Squalene Synthase Functional Motif" Joseph Chappell, PhD Pediatrics/Medical Genetics and Genomics - University of Alabama Med Ctr, Birmingham, AL Gregory J. Wehner, MD, PhD Biomedical Engineering "Validation, Optimiztion, and Image Processing of Spiral Cine Dense Magnetic Resonance Imaging for the Quantification of Left and Right Ventricular Mechanics" Brandon Fornwalt, MD, PhD Diagnostic Radiology/Research - Barnes-Jewish Hospital, St. Louis, MO Christopher Brown, MD, PhD Anatomy & Neurobiology "The Default Mode Network and Executive Function: Influence of Age, White Matter Connectivity, and Alzheimer's Pathology" Brian T. Gold, PhD Neurology - Hospital of the University of Pennsylvania, Philadelphia, PA Eseosa Ighodaro, MD, PhD Anatomy & Neurobiology "Studying Vascular Morphologies in the Aged Human Brain Using Large Autopsy Datasets" Peter Nelson, MD, PhD Neurology - Mayo Clinic School of Grad Med Educ, Rochester, MN
2019
Name Graduate Department Dissertation Title PhD Mentor Residency Kristin B. Linscott, MD, PhD Molecular and Cellular Biochemistry "Discovering a Novel Antifungal Target in Downstream Sterol Biosynthesis Using a Squalene Synthase Functional Motif" Joseph Chappell, PhD Pediatrics/Medical Genetics and Genomics - University of Alabama Med Ctr, Birmingham, AL Gregory J. Wehner, MD, PhD Biomedical Engineering "Validation, Optimiztion, and Image Processing of Spiral Cine Dense Magnetic Resonance Imaging for the Quantification of Left and Right Ventricular Mechanics" Brandon Fornwalt, MD, PhD Diagnostic Radiology/Research - Barnes-Jewish Hospital, St. Louis, MO Christopher Brown, MD, PhD Anatomy & Neurobiology "The Default Mode Network and Executive Function: Influence of Age, White Matter Connectivity, and Alzheimer's Pathology" Brian T. Gold, PhD Neurology - Hospital of the University of Pennsylvania, Philadelphia, PA Eseosa Ighodaro, MD, PhD Anatomy & Neurobiology "Studying Vascular Morphologies in the Aged Human Brain Using Large Autopsy Datasets" Peter Nelson, MD, PhD Neurology - Mayo Clinic School of Grad Med Educ, Rochester, MN

2018

Name Graduate Department Dissertation Title PhD Mentor Residency Erin Wolf Horrell, MD, PhD Physiology "Regulation of UV-Protective Pathways Downstream of the Melanocortin 1 Receptor in Melanocytes" John D'Orazio, MD, PhD General Surgery - Vanderbilt Univ Med Ctr, Nashville, TN Edita M. Klimyte, MD, PhD Molecular and Cellular Biochemistry "Elucidating Binding, Fusion, and Entry of Human Metapneumovirus" Rebecca E. Dutch, PhD Dermatology - University of Arkansas for Medical Sciences, Little Rock, AR
2018
Name Graduate Department Dissertation Title PhD Mentor Residency Erin Wolf Horrell, MD, PhD Physiology "Regulation of UV-Protective Pathways Downstream of the Melanocortin 1 Receptor in Melanocytes" John D'Orazio, MD, PhD General Surgery - Vanderbilt Univ Med Ctr, Nashville, TN Edita M. Klimyte, MD, PhD Molecular and Cellular Biochemistry "Elucidating Binding, Fusion, and Entry of Human Metapneumovirus" Rebecca E. Dutch, PhD Dermatology - University of Arkansas for Medical Sciences, Little Rock, AR

2017

Name Graduate Department Dissertation Title PhD Mentor Residency Lucas S. Broster, MD, PhD Behavioral Science "Emotional Enhancement and Repetition Effects During Working Memory in Persons with Mild Cognitive Impairment" Yang Jiang, PhD Psychiatry - University of California, San Francisco, CA
2017
Name Graduate Department Dissertation Title PhD Mentor Residency Lucas S. Broster, MD, PhD Behavioral Science "Emotional Enhancement and Repetition Effects During Working Memory in Persons with Mild Cognitive Impairment" Yang Jiang, PhD Psychiatry - University of California, San Francisco, CA

2016

Name Graduate Department Dissertation Title PhD Mentor Residency Dustin B. Stephens, MD, PhD Behavioral Science “Primary and secondary prevention of hepatitis C virus among rural Appalachian people who use drugs” Jennifer R. Havens, PhD, MPH Psychiatry - Harbor-UCLA Medical Center, Los Angeles, CA Dustin P. Brown, MD, PhD Anatomy & Neurobiology "Target-Directed Biosynthetic Evolution: Redirecting Plant Evolution to Genomically Optimize a Plant's Pharmacological Profile" Greg A. Gerhardt, PhD Radiology - UC San Diego, CA
2016
Name Graduate Department Dissertation Title PhD Mentor Residency Dustin B. Stephens, MD, PhD Behavioral Science “Primary and secondary prevention of hepatitis C virus among rural Appalachian people who use drugs” Jennifer R. Havens, PhD, MPH Psychiatry - Harbor-UCLA Medical Center, Los Angeles, CA Dustin P. Brown, MD, PhD Anatomy & Neurobiology "Target-Directed Biosynthetic Evolution: Redirecting Plant Evolution to Genomically Optimize a Plant's Pharmacological Profile" Greg A. Gerhardt, PhD Radiology - UC San Diego, CA

2014

Name Graduate Department Dissertation Title PhD Mentor Residency Andres Chang, MD, PhD Molecular and Cellular Biochemistry "Early Events of Human Metapneumovirus Infection" Rebecca E. Dutch, PhD Internal Medicine - Emory University, Atlanta, GA Mellani Lefta, MD, PhD Physiology     Pediatrics - Children's Hospital of the King's Daughters - Norfolk, VA Abdelghaffar Salous, MD, PhD Pharmacology "Mechanism of rapid elimination of lysophosphatidic acid and related lipids from the circulation of mice"   General Surgery - Sinai Hospital of Baltimore, Baltimore, MD
2014
Name Graduate Department Dissertation Title PhD Mentor Residency Andres Chang, MD, PhD Molecular and Cellular Biochemistry "Early Events of Human Metapneumovirus Infection" Rebecca E. Dutch, PhD Internal Medicine - Emory University, Atlanta, GA Mellani Lefta, MD, PhD Physiology     Pediatrics - Children's Hospital of the King's Daughters - Norfolk, VA Abdelghaffar Salous, MD, PhD Pharmacology "Mechanism of rapid elimination of lysophosphatidic acid and related lipids from the circulation of mice"   General Surgery - Sinai Hospital of Baltimore, Baltimore, MD

2013

Name Graduate Department Dissertation Title PhD Mentor Residency Zachary Fulkerson, MD, PhD Physiology "Lysophosphatidic Acid Production and Signaling in Platelets" Susan S. Smyth, MD, PhD Internal Medicine - Indiana University, Indianapolis, IN Carmen Kiper, MD, PhD Physiology "Strain-specific Cardiomyocyte Composition Regulates Cardiac Growth Potential" Jonathan Satin , PhD Pediatrics - University of Louisville, Kosair Children's Hospital, Louisville, KY Caitlin Latimer, MD, PhD Pharmacology   Nada Porter, PhD Anatomic Pathology - University of Washington Affiliated Hospital - Seattle, WA Scott Silva, MD, PhD Biochemistry "A Novel Role for VEGF Signialling in Carcinoid Tumors" B. Mark Evers, MD, FACS General Surgery Internship - Vanderbilt University Medical Center, Nashville, TN | Radiation Oncology Residency - Loyola University Medical Center, Chicago, IL Christopher Simmons, MD, PhD Physiology   Steven Estus, PhD Pediatrics - University of Kentucky, Lexington, KY
2013
Name Graduate Department Dissertation Title PhD Mentor Residency Zachary Fulkerson, MD, PhD Physiology "Lysophosphatidic Acid Production and Signaling in Platelets" Susan S. Smyth, MD, PhD Internal Medicine - Indiana University, Indianapolis, IN Carmen Kiper, MD, PhD Physiology "Strain-specific Cardiomyocyte Composition Regulates Cardiac Growth Potential" Jonathan Satin , PhD Pediatrics - University of Louisville, Kosair Children's Hospital, Louisville, KY Caitlin Latimer, MD, PhD Pharmacology   Nada Porter, PhD Anatomic Pathology - University of Washington Affiliated Hospital - Seattle, WA Scott Silva, MD, PhD Biochemistry "A Novel Role for VEGF Signialling in Carcinoid Tumors" B. Mark Evers, MD, FACS General Surgery Internship - Vanderbilt University Medical Center, Nashville, TN | Radiation Oncology Residency - Loyola University Medical Center, Chicago, IL Christopher Simmons, MD, PhD Physiology   Steven Estus, PhD Pediatrics - University of Kentucky, Lexington, KY

2012

Name Graduate Department Dissertation Title PhD Mentor Residency Natalie Wallis, MD, PhD Pharmacology   Michael W. Kilgore, PhD Pediatrics - University of Kentucky, Lexington, KY
2012
Name Graduate Department Dissertation Title PhD Mentor Residency Natalie Wallis, MD, PhD Pharmacology   Michael W. Kilgore, PhD Pediatrics - University of Kentucky, Lexington, KY

2011

Name Graduate Department Dissertation Title PhD Mentor Residency Romulo Albuquerque, MD, PhD Physiology/Ophthalmology "Soluble vascular endothelial growth factor receptor 2: an endogenous inhibitor of lymphangiogenesis" Jayakrishna Ambati, MD Ophthalmology - University of Kentucky, Lexington, KY James Butler, MD, PhD Microbiology & Immunology "Mechanisms governing the zonal activity of alpha-fetoprotein enhancer EIII" Brett Spear, PhD Psychiatry - UT Southwestern, Dallas, TX Willie Craft Jr., MD, PhD Molecular and Cellular Biochemistry "Investigation of the proteolytic processing of the hendra virus fusion protein" Rebecca E. Dutch, PhD Internal Medicine - Hahnemann University, Philadelphia, PA Michelle Stephens Humeidan, MD, PhD Anatomy & Neurobiology "Glutamate regulation in the hippocampal trisynaptic pathway in aging and status epilepticus" Greg Gerhardt, PhD Anesthesiology - Ohio State University, Columbus, OH Nirmal Ravi, MD, PhD Biomedical Engineering "Regeneration of damaged growth plate using IGF-I plasmid releasing porous PLGA scaffolds" David A. Puleo, PhD Family Medicine - Providence Hospital, Georgetown, Washington D.C.
2011
Name Graduate Department Dissertation Title PhD Mentor Residency Romulo Albuquerque, MD, PhD Physiology/Ophthalmology "Soluble vascular endothelial growth factor receptor 2: an endogenous inhibitor of lymphangiogenesis" Jayakrishna Ambati, MD Ophthalmology - University of Kentucky, Lexington, KY James Butler, MD, PhD Microbiology & Immunology "Mechanisms governing the zonal activity of alpha-fetoprotein enhancer EIII" Brett Spear, PhD Psychiatry - UT Southwestern, Dallas, TX Willie Craft Jr., MD, PhD Molecular and Cellular Biochemistry "Investigation of the proteolytic processing of the hendra virus fusion protein" Rebecca E. Dutch, PhD Internal Medicine - Hahnemann University, Philadelphia, PA Michelle Stephens Humeidan, MD, PhD Anatomy & Neurobiology "Glutamate regulation in the hippocampal trisynaptic pathway in aging and status epilepticus" Greg Gerhardt, PhD Anesthesiology - Ohio State University, Columbus, OH Nirmal Ravi, MD, PhD Biomedical Engineering "Regeneration of damaged growth plate using IGF-I plasmid releasing porous PLGA scaffolds" David A. Puleo, PhD Family Medicine - Providence Hospital, Georgetown, Washington D.C.

2010

Name Graduate Department Dissertation Title PhD Mentor Residency Karrie Grear, MD, PhD Physiology "Uncoupling protein-2 splicing efficiency and human disease" Steven Estus, PhD Neurology - Wake Forest University, Winston-Salem, NC Jeff Yates, MD, PhD Physiology "The genetic regulation of the response of hematopoietic stem/ progenitor cells to the cytostatic agent hydroxyurea" Gary Van Zant, PhD Internal Medicine & Pediatrics - Indiana University, Indianapolis, IN
2010
Name Graduate Department Dissertation Title PhD Mentor Residency Karrie Grear, MD, PhD Physiology "Uncoupling protein-2 splicing efficiency and human disease" Steven Estus, PhD Neurology - Wake Forest University, Winston-Salem, NC Jeff Yates, MD, PhD Physiology "The genetic regulation of the response of hematopoietic stem/ progenitor cells to the cytostatic agent hydroxyurea" Gary Van Zant, PhD Internal Medicine & Pediatrics - Indiana University, Indianapolis, IN

2009

Name Graduate Department Dissertation Title PhD Mentor Residency Janelle Reed, MD, PhD Anatomy & Neurobiology "Alterations in proteasome subunit expression and influences on the inflammatory response" Annadora J. Bruce-Keller, PhD Emergency Medicine - University of Kentucky, Lexington, KY
2009
Name Graduate Department Dissertation Title PhD Mentor Residency Janelle Reed, MD, PhD Anatomy & Neurobiology "Alterations in proteasome subunit expression and influences on the inflammatory response" Annadora J. Bruce-Keller, PhD Emergency Medicine - University of Kentucky, Lexington, KY

2008

Name Graduate Department Dissertation Title PhD Mentor Residency Mark A. Wurth, MD, PhD Biochemistry "Role of actin dynamics in paramyxovirus F mediated membrane fusion events" Rebecca E. Dutch, PhD Internal Medicine & Pediatrics - University of Kentucky, Lexington, KY Kristine S. Ziemba, MD, PhD Physiology "Targeting axon growth from neurons transplanted into the central nervous system" George M. Smith, PhD Neurology - Mayo Clinic, Pheonix, AZ
2008
Name Graduate Department Dissertation Title PhD Mentor Residency Mark A. Wurth, MD, PhD Biochemistry "Role of actin dynamics in paramyxovirus F mediated membrane fusion events" Rebecca E. Dutch, PhD Internal Medicine & Pediatrics - University of Kentucky, Lexington, KY Kristine S. Ziemba, MD, PhD Physiology "Targeting axon growth from neurons transplanted into the central nervous system" George M. Smith, PhD Neurology - Mayo Clinic, Pheonix, AZ

2007

Name Graduate Department Dissertation Title PhD Mentor Residency Brian K. Day, MD, PhD Anatomy & Neurobiology "Microelectrode array studies of normal and disease-altered L-glutamate regulation in the mammalian central nervous system" Greg Gerhardt, PhD Neurology - Washington University, St. Louis, MO
2007
Name Graduate Department Dissertation Title PhD Mentor Residency Brian K. Day, MD, PhD Anatomy & Neurobiology "Microelectrode array studies of normal and disease-altered L-glutamate regulation in the mammalian central nervous system" Greg Gerhardt, PhD Neurology - Washington University, St. Louis, MO

2006

Name Graduate Department Dissertation Title PhD Mentor Residency Michael W. Manning, MD, PhD Physiology "Differential effects of doxycycline, a broad spectrum inhibitor of matrix metalloproteinases" Alan Daugherty, PhD, DSc, FAHA Surgery Internship - University of Kentucky, Lexington, KY | Anesthesiology - University of Kentucky, Lexington, KY Erin L. Manning, MD, PhD Physiology "Regulation of telomere length in murine hematopoietic cells" Gary Van Zant, PhD Anesthesiology - University of Kentucky, Lexington, KY Xuan Nguyen, MD, PhD Anatomy & Neurobiology "Aging and the dynorphinergic system: evaluation of memory and motor systems in prodynorphin knockout mice" Guoying Bing, PhD Internal Medicine Internship - University of Cincinnati, Cincinnati, OH | Diagnostic Radiology Residency - University of Kentucky, Lexington, KY
2006
Name Graduate Department Dissertation Title PhD Mentor Residency Michael W. Manning, MD, PhD Physiology "Differential effects of doxycycline, a broad spectrum inhibitor of matrix metalloproteinases" Alan Daugherty, PhD, DSc, FAHA Surgery Internship - University of Kentucky, Lexington, KY | Anesthesiology - University of Kentucky, Lexington, KY Erin L. Manning, MD, PhD Physiology "Regulation of telomere length in murine hematopoietic cells" Gary Van Zant, PhD Anesthesiology - University of Kentucky, Lexington, KY Xuan Nguyen, MD, PhD Anatomy & Neurobiology "Aging and the dynorphinergic system: evaluation of memory and motor systems in prodynorphin knockout mice" Guoying Bing, PhD Internal Medicine Internship - University of Cincinnati, Cincinnati, OH | Diagnostic Radiology Residency - University of Kentucky, Lexington, KY

2005

Name Graduate Department Dissertation Title PhD Mentor Residency Christina S. Hines, MD, PhD Molecular and Cellular Biochemistry "X-ray crystallographic and functional studies on the neuropeptidase neurolysin" David W. Rodgers, PhD Psychiatry - Johns Hopkins University, Baltimore, MD
2005
Name Graduate Department Dissertation Title PhD Mentor Residency Christina S. Hines, MD, PhD Molecular and Cellular Biochemistry "X-ray crystallographic and functional studies on the neuropeptidase neurolysin" David W. Rodgers, PhD Psychiatry - Johns Hopkins University, Baltimore, MD

2004

Name Graduate Department Dissertation Title PhD Mentor Residency Joshua L. Hood, MD, PhD Microbiology & Immunology "The association of calpain and its regulatory proteins with the endoplasmic reticulum and golgi apparatus" Thomas L. Roszman, PhD Clinical Pathology - Washington University, St. Louis, MO Shane W. Rau, MD, PhD Physiology "Mechanisms of estradiol-mediated protection against stroke-like injury" Phyllis M. Wise, PhD Psychiatry - University of Virginia Health System, Charlottesville, VA
2004
Name Graduate Department Dissertation Title PhD Mentor Residency Joshua L. Hood, MD, PhD Microbiology & Immunology "The association of calpain and its regulatory proteins with the endoplasmic reticulum and golgi apparatus" Thomas L. Roszman, PhD Clinical Pathology - Washington University, St. Louis, MO Shane W. Rau, MD, PhD Physiology "Mechanisms of estradiol-mediated protection against stroke-like injury" Phyllis M. Wise, PhD Psychiatry - University of Virginia Health System, Charlottesville, VA

2003

Dena Dubal, MD, PhD

2003

Dena Dubal, MD, PhD

1999

Mathew A. Nicholls, MD, PhD

  • Dissertation: "MR studies of the acute response of the rhesus CNS to the neurotoxin MPTP"
    • Department of Biochemistry, Mentorship by Thomas C. Vanaman, PhD
  • Residency: Orthopedic Surgery - University of Kentucky, Lexington, KY
  • Current Position: Orthopedic Surgeon, Virginia Mason Franciscan Health, Bellevue, WA

Melisa W. Mullins, MD, PhD

  • Dissertation 
    • Department of Microbiology & Immunology, Mentorship by Charles Snow, PhD
  • Residency: Pediatrics - William Beaumont Hospital, Royal Oak, MI
  • Current Position: Pediatric Hematologist-Oncologist, Nationwide Children's Hospital Toledo, Toledo, OH
1999

Mathew A. Nicholls, MD, PhD

  • Dissertation: "MR studies of the acute response of the rhesus CNS to the neurotoxin MPTP"
    • Department of Biochemistry, Mentorship by Thomas C. Vanaman, PhD
  • Residency: Orthopedic Surgery - University of Kentucky, Lexington, KY
  • Current Position: Orthopedic Surgeon, Virginia Mason Franciscan Health, Bellevue, WA

Melisa W. Mullins, MD, PhD

  • Dissertation 
    • Department of Microbiology & Immunology, Mentorship by Charles Snow, PhD
  • Residency: Pediatrics - William Beaumont Hospital, Royal Oak, MI
  • Current Position: Pediatric Hematologist-Oncologist, Nationwide Children's Hospital Toledo, Toledo, OH

1994

Gregory E. Cooper, MD, PhD

  • Dissertation
    • Department of Microbiology & Immunology
  • Residency: Neurology - University of Iowa Hospitals & Clinics, Iowa City, IA
  • Current Position: Chief of Adult Neurology, Norton Healthcare; Director of Memory Center, Norton Neuroscience Institute, Louisville, KY
1994

Gregory E. Cooper, MD, PhD

  • Dissertation
    • Department of Microbiology & Immunology
  • Residency: Neurology - University of Iowa Hospitals & Clinics, Iowa City, IA
  • Current Position: Chief of Adult Neurology, Norton Healthcare; Director of Memory Center, Norton Neuroscience Institute, Louisville, KY

1992

Gregory N. Barnes, MD, PhD

1992

Gregory N. Barnes, MD, PhD